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| signature = | signature_alt = | website = | footnotes = | spouse = }} Robert Treat Paine (born 1933) is an ecologist and retired professor emeritus of zoology at The University of Washington, who coined the keystone species〔 concept in order to explain the relationship between ''Pisaster ochraceus'', a species of starfish, and ''Mytilus californianus'', a species of mussel.〔 〕 In his classic 1966 paper, Dr. Paine described such a system in Makah Bay in Washington State. This led to his 1969 paper where he proposed the keystone species concept. ==Early life and education== Paine grew up in Cambridge, Massachusetts, and attended Harvard University. After Harvard, Paine served in the U.S. Army where he was the battalion gardener. He later entered graduate school at the University of Michigan intending to study paleontology. After he took some courses in zoology and ecology at Michigan, his interests and studies changed. Upon graduating from the University of Michigan, Paine competed a post-doctoral fellowship at the Scripps Institution of Oceanography. In 1962, Paine joined the University of Washington where he spent the rest of his career and became well known for his work. 抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』 ■ウィキペディアで「Robert T. Paine (zoologist)」の詳細全文を読む スポンサード リンク
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